TODD, PETER ALAN
Associate Professor
Contact Information:
Department of Biological Sciences
National University of Singapore
14 Science Drive 4
Singapore 117543
6516 1034
6779 2486
dbspat@nus.edu.sg
Teaching Areas
LSM3254 – Ecology of Aquatic Environments
LSM4261 – Marine Biology
SP3202 – Aquatic Ecology Research
Research Areas
Marine biology, urban marine ecology, ecological engineering, coral reefs, animal colours, patterns and morphology.
Research Interests
I am an experimental marine ecologist who focusses on organism-environment interactions in nearshore waters, especially those close to urban centres. In both my curiosity-driven and translational work, I emphasise the design, build, implementation, and analysis of high-quality novel experiments. My research generates large quantities of new information and the great majority of my publications are data based. I am fundamentally concerned with increasing understanding of the ecology and functioning of tropical coastal marine organisms and communities.
Editorial Activities
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Editor-in-Chief for Oceanography and Marine Biology – An Annual Review (2022 to present)
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Editorial Board Member for Marine Ecology (2022 to present)
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Series Editor for Oceanography and Marine Biology – An Annual Review (2018 to 2022)
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Associate Editor [Zoology] for Nature in Singapore (2013 to present)
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Subject Editor [Marine Biology] for Oikos (2011 to present)
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Review Editor for Aquatic Biology (2007 to present)
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Associate Editor [Conservation and Ecology] for the Raffles Bulletin of Zoology (2005 to 2020)
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Academic Editor [Marine Biology] for PLoS ONE (2013 to 2018)
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Editorial Board Member for the Journal of Experimental Marine Biology and Ecology (2008 to 2016)
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International Advisory Board Member for the Singapore Journal of Tropical Geography (2007 to 2011)
Selected Publications
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Chang, C. C., and Todd, P. A. (2023). Reduced predation pressure as a potential driver of prey diversity and abundance in complex habitats. npj Biodiversity, 2(1), 1-5.
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Chan, S.H., Ong, D.R., Williams, G.A., Crickenberger, S., Loke, L.H., and Todd, P.A. (2022). Behaviour broadens thermal safety margins on artificial coastal defences in the tropics. Marine Environmental Research, 177, 105618
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Summers, S., Pek, Y. S., Vinod, D. P., McDougald, D., Todd, P.A., Birch, W. R., and Rice, S. A. (2022) Bacterial biofilm colonization and succession in tropical marine waters are similar across different types of stone materials used in seawall construction. Frontiers in Microbiology, 13
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Bauman, A.G., Januchowski-Hartley, F.A., Teo, A., and Todd, P.A. (2022) Further flattening of a degraded, turbid reef system following a severe coral bleaching event. Frontiers in Marine Science 9, 910085
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Ng, M.S., Teo, A., and Todd, P.A. (2022) Sediment trap height affects mass, particle size, and biogeochemical composition of material collected in an equatorial coral reef. Marine Pollution Bulletin, 183, 114086
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Omeyer, L.C., Duncan, E.M., Aiemsomboon, K., Beaumont, N., … Todd, P.A…. and Godley, B. J. (2022) Priorities to inform research on marine plastic pollution in Southeast Asia. Science of the Total Environment, 156704
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Yong, W.L.O., Todd, P.A., Ang, A.C., Ying, L.S., Teo, A., Lee, S.H., Low, J. and Neo, M.L. (2022) Fluted giant clam (Tridacna squamosa) restocking experiment in an urban turbid reef environment. Aquatic Conservation, 32: 633-644
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Nguyen, N.T.H, Friess, D.A. Todd, P.A., Mazor, T., Lovelock, C.E., Lowe, R., Gilmour, J., Huang, D. (2022) Maximising resilience to sea-level rise in urban coastal ecosystems through systematic conservation planning Landscape and Urban Planning, 221, 104374
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Hartanto, R.S., Loke, L.H. , Heery, E.C. , Hsiung, A.R. , Goh, M.W., Pek, Y.S., Birch, W.R. and Todd, P.A. † (2022). Material type weakly affects algal colonisation but not macrofaunal community in an artificial intertidal habitat. Ecological Engineering, 176, 106514.
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Goh, T.Z., Bauman, A.G., Januchowski-Hartley, F.A., Morgan, K. M., Seah, J.C. and Todd, P.A. (2021). Growth and carbonate production of crustose coralline algae on a degraded turbid reef system. Marine Pollution Bulletin, 173, 113135
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Mazor, T., Friess, D.A., Todd, P.A., Huang, D. Nguyen, N.T.H., Saunders, M., Runting, R.K., Lowe, R., Cartwright, P., Gilmour, J. and Lovelock, C.E. (2021) Large conservation opportunities exists in >90% of tropic-subtropic coastal habitats adjacent to cities. One Earth, 4: 1004-1015
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Ng, D., Daira, D. , Heery, E.C. and Todd, P.A. (2021) Antagonistic effects of seawalls and urban sedimentation on epilithic algal matrix (EAM)-feeding fishes. Marine Pollution Bulletin, 173, B113098
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Fong, J. and Todd, P.A. (2021) Spatio-temporal dynamics of coral–macroalgal interactions and their impacts on coral growth on urbanised reefs. Marine Pollution Bulletin, 172: 112849
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Mazor, T., Runting, R.K., Saunders, M.I., Huang, D., Friess, D.A., Nguyen, N.T., Lowe, R.J., Gilmour, J.P., Todd, P.A. and Lovelock, C.E., 2021. Future-proofing conservation priorities for sea level rise in coastal urban ecosystems. Biological Conservation, 260: 109190
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Pang, H.E., Poquita-Du, R.C, Jain, S.S., Huang, D. and Todd, P.A. (2021) Among-genotype responses of corals to emersion: implications for eco-engineering seawalls. Marine Environmental Research, 168: 105312
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Seah, J.C.L., Bauman G. and Todd, P.A. (2021) Temporal variation in macroalgal removal: insights from an impacted equatorial coral reef system. Marine Biology, 168: 1-12
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Chan, I.Z., Wang, W.Y. and Todd, P.A. (2021). Facial band colour in the mangrove crab Parasesarma peninsulare Shahdadi, Ng and Schubart, 2019 plays a role in mate recognition. Estuarine, Coastal and Shelf Science, 106721
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Prasetia, R., Lim, Z.W., Teo, A., Shlesinger, T., Loya, Y. and Todd, P.A. (2020) Population dynamics and growth rates of free-living mushroom corals (Scleractinia: Fungiidae) in the sediment-stressed reefs of Singapore. Advances in Marine Biology, 87: 115-140.
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Taira, D., Heery, E.C., Loke, L.H.L., Teo, A., Bauman, A.G. and Todd, P.A. (2020) Ecological engineering across organismal scales: trophic-mediated positive effects of microhabitat enhancement on fish. Marine Ecology Progress Series. org/10.3354/meps13462
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Poquita-Du, R.C., Goh, Y.L, Chou, L.M. and Todd, P.A. (2020) Gene expression and photophysiological changes in Pocillopora acuta coral holobiont following heat stress and recovery. Microorganisms, 8: 1227
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Seng, N., Lai, S., Fong, J., Faiq, M., Cheng, C., Cheok Z.Y. and Todd P.A. (2020) Early evidence of microplastics on seagrass and macroalgae. Marine and Freshwater Research 71, 922-928
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Hsiung, A.R., Tan, W.T., Loke, L.H.L., Firth, L.B., Heery, E.C., Ducker, J., Clark, V., Pek, Y.S., Birch, W.R., Ang, A.C.F., Hartanto, R.S., Chai, T.M.F. and Todd P.A. (2020) Little evidence that lowering the pH of concrete supports greater biodiversity on tropical and temperate seawalls. Marine Ecology Progress Series, 656, 193-205.
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Fong, J., Deignan, L.K., Bauman, A.G., Steinberg, P.D., McDougald, D and Todd P.A. (2020). Contact-and water-mediated effects of macroalgae on the physiology and microbiome of three Indo-Pacific coral species. Frontiers in Marine Science, 6: 831
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Smith, E., Gurskaya, A., Hume, B., Voolstra, C., Todd P.A., Bauman, A.G. and Burt, J. (2020) Low Symbiodiniaceae diversity in a turbid marginal reef environment. Coral Reefs, 39: 545-553
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Heery, E.C., Lian, K.Y., Loke, L.H.L., Tan, H.T. and Todd P.A. (2020). Evaluating seaweed farming as an eco-engineering strategy for ‘blue’ shoreline infrastructure. Ecological Engineering, 152: 105857
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Heery, E.C., Oh, R.K., Taira, D., Ng, D., Kong, C.C., Hartanto, R.S., Hsiung, A.R., Chai, T.M., Loke, L.H.L., Yeo, H.H. and Todd P.A. (2020). Human-engineered hydrodynamic regimes as a driver of cryptic microinvertebrate assemblages on urban artificial shorelines. Science of The Total Environment, 138348
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Januchowski-Hartley, F.A., Bauman, A.G., Morgan, K.M., Seah, J.C.^, Huang, D. and Todd P.A. (2020). Accreting coral reefs in a highly urbanized environment. Coral Reefs, 39: 717–731
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Monteil, Y., Teo, A., Fong, J., Bauman, A.G. and Todd, P.A. (2020). Effects of macroalgae on coral fecundity in a degraded coral reef system. Marine Pollution Bulletin, 151: 11089
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Poquita-Du, R.C., Huang, D., Chou, L.M. and Todd, P.A. (2020). The contribution of stress-tolerant endosymbiotic dinoflagellate Durusdinium to Pocillopora acuta survival in a highly urbanized reef system. Coral Reefs, 39: 745–755
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Kong, E., Ow, Y.X., Lai, S., Yaakub, S.M. and Todd P.A. (2020) Effects of shading on seagrass morphology and thermal optimal of productivity. Marine and Freshwater Research, 71, 913-921.
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Yaakub, S.M., Foster, N.B., Waycott, M. and Todd, P.A. (2020) Translating seagrass science into action. Marine and Freshwater Research, 71: i-iii
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Todd, P.A., Heery, E.C., Loke, H.L., Thurstan, R.H., Kotze J. and Swan, C. (2019) Towards an urban marine ecology: characterizing the drivers, patterns and processes of marine ecosystems in coastal cities. Oikos, 128: 1215-1242
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Poquita-Du, R.C., Quek, R., Jain, S.S., Schmidt-Roach, S., Tun, K., Heery, E,C., Chou, L.M., Todd, P.A. and Huang, D. (2019) Last species standing: loss of Pocilloporidae corals associated with coastal urbanization in a tropical city state. Marine Biodiversity, 49: 1727–1741
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Bauman, A.G., Seah, J.C., Januchowski-Hartley, F.A., Hoey, A.S., Fong, J. and Todd P.A. (2019). Fear effects associated with predator presence and habitat structure interact to alter herbivory on coral reefs. Biology Letters, 15: 20190409
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Lim, A.Y., Chan, I.Z., Carrasco, L.R. and Todd, P.A. (2019). Aposematism in pink warty sea cucumbers: independent effects of chromatic and achromatic cues. Marine Ecology Progress Series 631: 157-164
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Morris, R.L., Heery, E.C., Loke, L.H.L., Lau E., Strain, E.M.A., , Airoldi, L., Alexander, K.A., Bishop, M.J., Coleman, R.A., Cordell, J.R., Dong, Y-W., . Firth, L.B., Hawkins, S.J., Heath, T., Kokora, M., Lee, S.Y., Miller, J.K., Perkol-Finke, S., Rella, A., Steinberg, P.D., Takeuchi, I., Thompson, R.C., Todd, P.A., Toft, J.D. and Leung, K.M.Y. (2019) Design options, implementation issues and evaluating success of ecologically-engineered shorelines. Oceanography and Marine Biology: An Annual Review, 55: 87-388
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Poquita-Du, R.C. , Huang, D., Chou, L.M., Mrinalini and Todd P.A. (2019) Short term exposure to heat and sediment triggers changes in coral gene expression and photo-physiological performance. Frontiers in Marine Science, 6: 121
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Loke, H.L., Chisholm, R.A. and Todd P.A. (2019) Effects of habitat area and spatial configuration on biodiversity in an experimental intertidal community at a landscape scale. Ecology, 100, e02757
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Chan, I.Z.W. , Stevens, M. and Todd P.A. (2019) PAT-GEOM: A software package for the analysis of animal colour pattern geometry10: 591–600
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Loke, H.L., Heery, E.C., Lai, S., Bouma, T.J. and Todd, P.A. (2019) Area-independent effects of water-retaining features on intertidal biodiversity on eco-engineered seawalls in the tropics. Frontiers in Marine Science, 6:16
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Teo, A. and Todd, P.A. (2018) Simulating the effects of colony density and intercolonial distance on fertilisation success in broadcast spawning scleractinian corals. Coral Reefs, 37: 891-900
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Heery, E.C., Hoeksema, B.W., Browne, N.K., Reimer, J.D., Ang, P.O., Huang, D., Friess, D.A., Chou, L.M., Loke, L.H.L., Saksena-Taylor, P., Alsagoff, N., Yeemin, T., Sutthacheep, M., Vo, S.T., Bos, A.R., Gumanao, G.S., Hussein, M.A.S., Waheed, Z., Lane, D.J.W., Johan, O., Kunzmann, A., Jompa, J., Suharsono, Taira, D., Bauman, A.G. and Todd, P.A. (2018) Urban coral reefs: Degradation and resilience of hard coral assemblages in coastal cities of East and Southeast Asia. Marine Pollution Bulletin, 135:654-681
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Tan, W.T., Loke, L.H.L., Yeo, D.C.J., Tan, S.K. and Todd, P.A. (2018) Do Singapore’s seawalls host non-native marine molluscs? Aquatic Invasions 13:365–378
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Sew, G., Calbet, A., Drillet, G. and Todd, P.A. (2018) Effects of concentration and size of suspended particles on the ingestion, reproduction and mortality rates of the copepod, Acartia tonsa. Marine Environmental Research 140: 251-264
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Sim, D.Z., Neo, M.L., Ang, A.C., Ying, L.S. and Todd P.A. (2018) Trade-offs between defence and competition in gregarious juvenile fluted giant clams (Tridacna squamosa L.). Marine Biology 165:103 https://doi.org/10.1007/s00227-018-3363-6
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Lai, S., Loke, L.H., Bouma, T.J. and Todd, P.A. (2018) Biodiversity surveys and stable isotope analyses reveal key differences in intertidal assemblages between tropical seawalls and rocky shores. Marine Ecology Progress Series 587:41-53
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Bauman, A.G. , Hoey, A.S., Dunshea, G., Feary, D.A., Low, J. and Todd P.A. (2017) Macroalgal browsing on a heavily degraded, urbanized equatorial reef system. Scientific Reports 7(1) Article number 8352
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Browne, N.K., Yaakub, S.M., Tay, J.K.L. and Todd P.A. (2017) Recreating the shading effects of ship wake induced turbidity to test acclimation responses in the seagrass Thalassia hemprichii. Estuarine, Coastal and Shelf Science 199: 87-95
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Neo, M.L., Wabnitz, C.C.C., Braley, R.D., Heslinga, G.A., Fauvelot, C., Van Wynsberge, S., Andréfouët, S., Waters, C., Tan A. S.-H., Gomez, E.D., Costello, M.J. and Todd P.A. (2017) Giant clams (Bivalvia: Cardiidae: Tridacninae): A comprehensive update of species and their distribution, current threats and conservation status. Oceanography and Marine Biology: An Annual Review 55:87-387
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Larsen, T.C., Browne, N.K. , Erichsen, A.C., Tun, K. and Todd P.A. (2017) Modelling for management: Coral photo-physiology and growth potential under varying turbidity regimes. Ecological Modelling 362:1-12
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Toh, K.B. and Todd, P.A. (2017) Camouflage that is spot on! Optimization of spot size in prey-background matching. Evolutionary Ecology 31:447-461
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Tanzil J.T.I., Lee J.N., Brown B.E., Quax R., Kaandorp J.A., Lough J.M. and Todd P.A. (2016) Luminescence and density banding patterns in massive Porites corals around the Thai-Malay Peninsula, Southeast Asia. Limnology and Oceanography 61: 2003-2026
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Teo. A., Guest, J.R., Neo, M.L., Vicentuan-Cabaitan, K., and Todd, P.A. (2016) Quantification of coral sperm collected during a synchronous spawning event. PeerJ 4 e2180
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Guest J.R., Low, J., Tun, K., Wilson, B., Ng, C., Raingeard, D., Ulstrup, K.E., Tanzil, J.T.I.#, Todd, P.A., Toh, T.C., McDouglad, D., Chou L.M. and Steinberg, P.D. (2016) Coral community bleaching response on a highly urbanised reef. Scientific reports6, 20717; doi: 10.1038/ srep20717
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Loke L.H.L. and Todd, P.A. (2016) Structural complexity and component type increase intertidal biodiversity independently of area. Ecology97: 383–393
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Todd, P. A. , Phua, H., and Toh, K.B. (2015) Interactions between background matching and disruptive colouration: Experiments using human predators and virtual crabs. Current Zoology 61: 718–728
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Bauman, A.G., Guest J.R., Dunshea, G., Low, J., Todd, P.A., and Steinberg, P.D. (2015) Coral settlement on a highly disturbed equatorial reef system. PLOS ONE10: e0127874. doi:10.1371/journal.pone.0127874
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Neo, M.L., Vicentuan-Cabaitan, K., Teo, S. L.-M., Erftemeijer, P.L.A. and Todd, P.A. (2015) Early life history of the fluted giant clam, Tridacna squamosa and its potential effects on larval dispersal models. Journal of Experimental Marine Biology and Ecology 469: 76-82
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Browne, N.K., Tay, J.K.L., Low, J., Larson, O. and Todd, P.A. (2015) Fluctuations in coral health of four common inshore reef corals in response to seasonal and anthropogenic changes in water quality. Marine Environmental Research. 105: 39-52
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Loke L.H.L., Ladle, R.J., Bouma, T.J. and Todd, P.A. (2015) Creating complex habitats for restoration and reconciliation. Ecological Engineering. 77: 307-313
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Neo, M.L., Eckman, W., Vicentuan-Cabaitan, K., Teo, S. L.-M. and Todd, P.A. (2015) The ecological significance of giant clams in coral reef ecosystems. Biological Conservation. 181: 111–123
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Junjie, R.K, Browne, N.K., Erftemeijer P.L.A. and Todd, P.A. (2014) Impacts of sediments on coral energetics: partitioning the effects of turbidity and settling particles. PLOS ONE.9(9): e107195. doi:10.1371/journal.pone.0107195
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Soo, P. and Todd, P.A. (2014) The behaviour of giant clams (Bivalvia: Cardiidae: Tridacninae). Marine Biology.161:2699–2717
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Tay, Y.C., Noreen, A.M.E., Chou, L.M. and Todd, P.A. (2014) Connectivity of the broadcast spawning reef coral Platygyra sinensis on impacted reefs, and the description of new microsatellite markers. Coral Reefs.10.1007/s00338-014-1206-6
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Lee, B.Y., Parra-Velandia, F.J., Ng, N.K., and Todd, P.A. (2014) An unusual form of camouflage in the mangrove crab Clistocoeloma merguiense. Bulletin of Marine Science. 90: 967-968
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Yaakub, S.M., Chen, E., Bouma, T.J., Erftemeijer, P.L.A. and Todd, P.A. (2014) Chronic light reduction reduces overall resilience to additional shading stress in the seagrass Halophila ovalis. Marine Pollution Bulletin. 83: 467–474
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Yaakub, S.M., McKenzie, L.J., Erftemeijer P.L.A., Bouma, T.J. and Todd, P.A. (2014) Courage under fire: Seagrass persistence adjacent to a highly urbanised city-state. Marine Pollution Bulletin. 83: 417–424
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Browne, N.K., Precht, E., Last, K.S. and Todd, P.A. (2014) The photo-physiological costs associated with acute sediment stress events in three near-shore turbid water corals. Marine Ecology Progress Series. 502:129-143
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Loke L.H.L., Jachowski, N.R., Bouma, T.J., Ladle, R.J. and Todd, P.A. (2014) Complexity for Artificial Substrates (CASU): software for creating and visualising habitat complexity. PLOS ONE 9(2): e87990. doi:10.1371/ journal.pone.0087990
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Neo, M.L., Erftemeijer, P.L.A., van Beek J.K.L., van Maren D. S., Teo S. L-M and Todd, P.A. (2013) Recruitment constraints in Singapore’s fluted giant clam (Tridacna squamosa) populations – A dispersal model approach. PLOS ONE. 8(3): e58819. DOI:10.1371/journal.pone.0058819
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Tanzil, J.T.I., Brown, B.E., Dunne, R.P., Lee, J.N., Kaandorp, J.A. and Todd, P.A. (2013) Regional decline in growth rates of massive Porites corals in Southeast Asia. Global Change Biology. 19: 3011-3023
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Stevens, M., Cheo P.R. and Todd, P.A. (2013) Color change and camouflage in the horned ghost crab, Ocypode ceratophthalmus. Biological Journal of the Linnean Society. DOI: 10.1111/bij.12039
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Ong, C.W., Reimer, J.D. and Todd, P.A. (2012) Morphologically plastic responses to shading in the zoanthids Zoanthus sansibaricus and Palythoa tuberculosa. Marine Biology. 160:1053–1064
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Neo, M.L. and Todd, P.A. (2012) Population density and genetic structure of the giant clams Tridacna crocea and T. squamosa on Singapore’s reefs. Aquatic Biology. 14: 265–275
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Todd, P.A., Oh, J., Loke L.H.L., and Ladle, R.J (2012) Multi-scale phenotype-substrate matching: evidence from shore crabs (Carcinus maenas L.).Ecological Complexity. 12: 58-62
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Erftemeijer, P.L.A., Riegl B., Hoeksema, B.W. and Todd, P.A. (2012) Environmental impacts of dredging and other sediment disturbances on corals: a review. Marine Pollution Bulletin. 64: 1737–1765
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Tay, Y.C., Todd, P.A., Per, S.R. and Chou, L.M. (2012) Simulating the transport of coral larvae among the Southern Islands of Singapore.Aquatic Biology. 15: 283-297
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Todd, P.A., Wang W.Y., Huang, H. Belle, C.C., Lim .M L.M., Yeo, D. C. J. (2011) The function of colourful facial bands in mangrove crab (Perisesarma) communication. Journal of Experimental Marine Biology and Ecology. 407: 26–33
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Belle, C., Wong, J.Q.H., Yeo, D.C.J., Tan, S.H.,Tan, H.H., Clews, E., Todd, P.A. (2011) Ornamental trade as a pathway of Cherax quadricarinatus introduction and establishment. Aquatic Biology. 12: 69-79
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Tay, Y.C., Guest, J.R., Todd, P.A. and Chou, L.M. (2011) Vertical distribution and settlement competencies in broadcast spawning coral larvae: implications for dispersal. Journal of Experimental Marine Biology and Ecology. 409: 324–330
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Neo, M.L. and Todd, P.A. (2010) Predator-induced changes in fluted giant clam (Tridacna squamosa L.) shell morphology. Journal of Experimental Marine Biology and Ecology. 397: 21–26
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Todd, P.A., Ong, X. and Chou, L.M. (2010) Impacts of pollution on marine life in Southeast Asia. Biodiversity and Conservation. 19: 1063-1082
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Ow Y.X. and Todd, P.A. (2010) Light-induced morphological plasticity in the Scleractinian coral Goniastrea pectinata and its functional significance. Coral Reefs. 29: 797-808
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Todd, P.A., Guest, J.R., Lu, J. and Chou, L.M. (2010) One in four citations in marine biology papers are inappropriate. Marine Ecology Progress Series. 408: 299-303
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Huang, D., Meier, R., Todd, P.A. and Chou, L.M. (2009) More evidence for pervasive paraphyly in scleractinian corals: systematic study of Southeast Asian Faviidae (Cnidaria; Scleractinia) based on molecular and morphological data. Molecular Phylogenetics and Evolution. 50: 102-116
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Reimer, J.D. and Todd, P.A. (2009) Preliminary molecular examination of zooxanthellate zoanthid (Hexacorallia: Zoantharia) and associated zooxanthellae (Symbiodinium spp.) diversity in Singapore. Raffles Bulletin of Zoology. 22: 103-120 [partial front cover]
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Todd, P.A., Lee J. H. and Chou L.M. (2009) Polymorphism and crypsis in the boring clam (Tridacna crocea): potential strategies against visual predators. Hydrobiologia. 635: 37-43 [front cover]
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Todd, P.A. (2008) Morphological plasticity in scleractinian corals. BiologicalReviews. 83: 315-337 [front cover]
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Han, L., Todd, P.A., Chou, L.M., Yap, V.B. and Sivalonganathan, B. (2008) The defensive role of scutes in juvenile fluted giant clams (Tridacna squamosa). Journal of Experimental Marine Biology and Ecology. 359: 77-83
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Todd, P.A. and Ladle, R.J. (2008) Citations: poor practices by authors reduce their value. Nature. 451: 244
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Huang, D., Meier, R., Todd, P.A. and Chou, L.M. (2008) Slow mitochondrial COI sequence evolution at the base of the metazoan tree and its implications for DNA barcoding. Journal of Molecular Evolution. 66:167-174
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Todd, P.A., Yeo, D.C.J., Li, D. and Ladle, R.J. (2007) Citing practices in ecology: can we believe our own words? Oikos. 116: 1599-1601
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Huang, D., Todd, P.A. and Guest, J.R. (2007) Locomotion and aggregation in the fluted giant clam (Tridacna squamosa L.). Journal of Experimental Marine Biology and Ecology. 342: 269–281.
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Todd, P.A., Briers, R.A., Ladle, R.J., Middleton, F. (2006) Phenotype-environment matching in the shore crab (Carcinus maenas). Marine Biology. 148: 1357-1367.
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Todd, P.A. and Chou, L.M. (2005) A Tale of Survival: Labrador Park, Singapore. Coral Reefs. 24 (3): 391-391.
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Todd, P.A., Ladle, R.J., Briers, R. and Brunton, A. (2005) Quantifying two-dimensional dichromatic patterns using a photographic technique: case study on the shore crab (Carcinus maenas, L.). Ecological Research. 20: 497-502.
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Todd, P.A., Ladle, R.J., Lewin-Koh, N.J.I. and Chou, L.M. (2004) Flesh or Bone? Quantifying small-scale coral morphology using with-tissue and without-tissue techniques. Marine Biology.145: 323-328.
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Todd, P.A., Sidle, R.C. and Lewin-Koh, N.J.I. (2004) An aquarium experiment for identifying the physical factors inducing morphological change in two massive scleractinian corals. Journal of Experimental Marine Biology and Ecology. 299: 97-133.
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Todd, P.A., Ladle, R.J., Lewin-Koh, N.J.I. and Chou, L.M. (2004) Genotype x environment interactions in transplanted clones of the massive corals Favia speciosa and Diploastrea heliopora. Marine Ecology Progress Series. 271: 167-182.
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Todd, P.A., Sidle, R.C. and Chou, L.M. (2002) Plastic corals from Singapore 2. Coral Reefs. 21: 407-408.
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Todd, P.A., Sidle, R.C. and Chou, L.M. (2002) Plastic corals from Singapore 1. Coral Reefs. 21: 391-392.
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Todd, P.A., Sanderson, P.G. and Chou, L.M. (2001) Small-scale variation in the scleractinian coral Favia speciosa (Dana, 1846) around Singapore. Hydrobiologia. 444: 227-235